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Code: 83795
ECTS: 5.0
Lecturers in charge: prof. dr. sc. Goran Kovačević
Lecturers: prof. dr. sc. Goran Kovačević - Seminar

dr. sc. Maja Novosel - Practicum
Take exam: Studomat
English level:


All teaching activities will be held in Croatian. However, foreign students in mixed groups will have the opportunity to attend additional office hours with the lecturer and teaching assistants in English to help master the course materials. Additionally, the lecturer will refer foreign students to the corresponding literature in English, as well as give them the possibility of taking the associated exams in English.

1. komponenta

Lecture typeTotal
Lectures 30
Practicum 30
Seminar 15
* Load is given in academic hour (1 academic hour = 45 minutes)
Recognize and undestand evolutionary subjects in environmental sciences.
Knowledge of the basic evolutionary mechanisms in biology.
Compare the historical context and modern evolutionary science.
Apply mathematical models in phylogeny and evolution.
Analyze scientific and popular papers from the field of evolution.
Present the working results by oral, written and multimedial presentation.

1. Lectures: Evolution as a scientific discipline and its historical development
Seminar: Researchers and scientists in evolution
Practical work: Microfossils, Collections

2. Lectures: Cosmic evolution, Solar system and formation of Earth
Seminar: Objects in space
Practical work: Macrofossils

3. Lectures: Chemical evolution, pre-atmosphere and conditions on Earth
Seminar: Hydrothermal vents
Practical work: Microfossils preparations and electronic map

4. Lectures: Biological evolution; Domains and kingdoms of the living world; Universal ancestors of the living beings
Seminar: Adaptations and biodiversity
Practical work: Coacervates, literature analysis

5. Lectures: Evolution of eukaryotic cell and organelles
Seminar: Complex plastids and nucleomorph
Practical work: Macromolecules of biological importance

6. Lectures: Symbiosis and endosymbiosis; The role of symbiosis in speciation and biodiversity; Evolutionary models of symbiosis
Seminar: Humans as symbiotic system
Practical work: Examples of symbiosis

7. Lectures: Green hydra "living fossil"
Seminar: Protective coloration
Practical work: Green hydra, symbiotic model

8.-9. Lectures: Living world in the past, Evidences of evolution
Seminar: Evolutionary motivs in literature and on film, Fossils in Croatia
Practical work: Isolation and culturing of endosymbionts

10. Lectures: Emergence of Metazoa; Molecular evolution and molecular phylogeny
Seminar: Cambrian explosion
Practical work: Microcosm models; adaptations

11. Lectures: Microevolution, macroevolution and megaevolution
Seminar: Genetic variability
Practical work: Hardy-Weinberg law, variability

12.-13. Lectures: Population genetics
Seminar: Migrations in human evolution
Practical work: Genetic drift, migrations

14.-15. Lectures: Human evolution, Hominids, Prehistoric cultures
Seminar: Paleoanthropological sites in Krapina and Vindija
Practical work: Mutations

Possible visits: Archaeological Museum in Zagreb, ZOO, Rokov Perivoj Park
  1. Stearns, S. C., Hoekstra, R. F. 2005. Evolution an Introduction. 2nd Edition Oxford University Press.
  2. Kovačević, G. Praktikumski priručnik Simbioza hidre i alge, 2013. (za internu uporabu).
  3. Lewin, R. 2005. Human Evolution - An Illustrated Introduction, Blackwell Publishing Ltd.
6. semester
Mandatory course - Regular study - Environmental Sciences
Consultations schedule:
  • prof. dr. sc. Goran Kovačević:

    After each lecture and seminar and upon e-mail contact.

  • prof. dr. sc. Goran Kovačević:

    After each lecture and seminar and upon e-mail contact.